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The glans extends about halfway down the shaft and is spiny except at the tip.In comparison, the glans of felids is short and spiny, while that of viverrids is smooth and long.The male waterfowl evolution of a phallus to forcefully copulate with females has led to counteradaptations in females in the form of vaginal structures called dead end sacs and clockwise coils.These structures make it harder for males to achieve intromission.Fossa mating includes a copulatory tie, which may be enforced by the male's spiny penis.This unusually lengthy mating is due to the physical nature of the male's erect penis, which has backwards-pointing spines along most of its length. abramus is more curved, the shaft and the tip are more robust in P.

Even within the Vertebrata there are morphological variants with specific terminology, such as hemipenes.Male and female emus are similar in appearance, The male tinamou has a corkscrew shaped penis, similar to those of the ratites and to the hemipenis of some reptiles.Females have a small phallic organ in the cloaca which becomes larger during the breeding season.The lake duck is notable for possessing, in relation to body length, the longest penis of all vertebrates; the penis, which is typically coiled up in flaccid state, can reach about the same length as the animal himself when fully erect, but is more commonly about half the bird's length.It is theorized that the remarkable size of their spiny penises with bristled tips may have evolved in response to competitive pressure in these highly promiscuous birds, removing sperm from previous matings in the manner of a bottle brush.

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